Crossword Genius

Pathogen that turns up regularly in gill pouches (1,4)


I believe the answer is:

e coli

'pathogen' is the definition.
(E coli is a pathogenic bacterium)

'turns up regularly in gill pouches' is the wordplay.
'turns up' is a reversal indicator.
'regularly in' indicates alternate letters.
The alternating letters of 'gillpouches' are 'iloce'.
'iloce' back-to-front is 'E COLI'.

'that' is the link.

(Other definitions for e coli that I've seen before include "Cause of internal problem" , "cause of illness" , "cause of serious complaint" , "cause of ill-feeling" , "An infection found in food and water" .)

I've seen this clue in The Guardian.
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