They were fabulously sour to the fox (6)
I believe the answer is:
grapes
'they' is the definition.
Both the answer and definition are plural nouns.
Perhaps they are linked in a way I don't understand?
I don't understand the rest of the clue.
Can you help me to learn more?
(Other definitions for grapes that I've seen before include "Fruit that grows in clusters" , "Vine fruits" , "See 10 Across" , "Vine fruit" , "They grow on vines" .)