Crossword Genius

Predator one across upset (4)

Ross

I believe the answer is:

wolf

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'predator' is the definition.

I don't understand how the remainder of the clue works.

Can you help me to learn more?

(Other definitions for wolf that I've seen before include "Howling canine that hunts in packs" , "Carnivore -- breed of 13 down" , "Devour ravenously" , "Pack animal; composer" , "gorge" .)

I've seen this clue in The Times.
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